Mark-recapture (Lincoln) estimate

The Mark-recapture (Lincoln) estimate computes mark-recapture (lincoln) estimate from the relation N = M C / R. It takes 3 inputs (marked, second catch, recaptured) and returns the population. It is written for biology students, laboratory technicians and field researchers who want a quick, repeatable figure when planning a culture or a dilution series, working through a growth, inheritance or sequence problem, or comparing organisms and populations of different sizes. Enter your values below and the result updates instantly, and you can share a permalink that pre-fills the exact calculation. Keep your units consistent across every input, using a single time unit wherever a rate or an interval appears and matching length, area and volume units wherever a ratio is involved, enter measured values rather than rounded ones so that any rounding happens only at the end, and when a count comes from a sample rather than the whole, such as a hemocytometer field or a survey plot, keep track of the factor that scales it back up. For example, with marked = 100, second catch = 100, recaptured = 20, the population works out to 500, and the worked example further down the page shows every step so you can follow the arithmetic and reproduce it by hand. This is a standard formula rather than a jurisdiction-specific rule, so it does not depend on an external rate or schedule, and the figure above each result carries the date it was last verified. The figure follows from the numbers you enter, and living material rarely holds that still: growth slows as nutrients run down and waste accumulates, rates shift with temperature, and individuals vary within any population, so treat the result as a working figure and expect what you see at the bench or in the field to sit near it rather than exactly on it.

With Marked = 100, Second catch = 100, Recaptured = 20, the result is 500.

Formula: N = M C / R. Source: our documented method, as at 2026-06-22.

Population500

Applies to: any numeric inputs. Method: a standard formula, not a jurisdiction-specific rule. How we document and test our methods, checked 2026-06-22.

The formula

N = M C / R

Worked example

With Marked = 100, Second catch = 100, Recaptured = 20:

  1. N = M C / R
  2. Population = 500

This worked example is one of the automated golden-value tests this calculator must pass before it can publish.

What this assumes

  • Inputs are real numbers in the units shown.
  • The result is the exact value of N = M C / R; general information, not professional advice.

Frequently asked questions

What formula does this use?

N = M C / R. This is a standard formula rather than a jurisdiction-specific rule.

Does the result ever change over time?

No. This is a pure formula with no external rate, so the same inputs always give the same result.

Official sources and verification

Reviewed by the CalculatorHub team, edited by James Graham, 2026-06-22. See our methodology. General information, not professional advice.